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Having the proper musical pitch. People don't realize how hard it is to sing in tune when you can't hear yourself!
Farlex Dictionary of Idioms. © 2015 Farlex, Inc, all rights reserved.
in a state where musical notes are at their proper intervals so that none are flat or sharp. Your piano is in tune. The choir members all sang in tune.
McGraw-Hill Dictionary of American Idioms and Phrasal Verbs. © 2002 by The McGraw-Hill Companies, Inc.
Also, in tune with.
1. In agreement in musical pitch or intonation, as in It's hard to keep a violin in tune during damp weather, or Dave is always in tune with the other instrumentalists. [Mid-1400s]
2. In concord or agreement, as in He was in tune with the times. [Late 1500s] The antonyms for both usages, dating from the same periods, are not in tune and out of tune, as in That trumpet's not in tune with the organ, or The lawyer was out of tune with his partners.
The American Heritage® Dictionary of Idioms by Christine Ammer. Copyright © 2003, 1997 by The Christine Ammer 1992 Trust. Published by Houghton Mifflin Harcourt Publishing Company. All rights reserved.