politum, because the male copulates with the female not only when she is building the mud tube but also when she is provisioning it or building a cell partition.
According to Alcock (1991), when a female copulates more than once, she enables the competition between the sperm of her partners.
Among gorillas, for example, one female copulates
with one male during a given fertile period.
Second, a lone large male sometimes foregoes the fighting, guarding, and pairing phases and quickly grabs a paired female and copulates with her; this behavior has been recorded when the female is very close to the egg beds (Video 1).
This lower copulatory success for unpaired males might reflect female choice; indeed, we have observed females jet from males that attempt to copulate in the field and laboratory.
Resident, related females in the new troops retain an edge over the incoming males in controlling who eats what and who copulates