According to Alcock (1991), when a female copulates more than once, she enables the competition between the sperm of her partners.
Once the female returns with mud or spiders, the satellite male quickly jumps her trying to copulate.
Among gorillas, for example, one female copulates
with one male during a given fertile period.
Second, a lone large male sometimes foregoes the fighting, guarding, and pairing phases and quickly grabs a paired female and copulates
with her; this behavior has been recorded when the female is very close to the egg beds (Video 1).
Resident, related females in the new troops retain an edge over the incoming males in controlling who eats what and who copulates
The loyal angels copulate
in their special way but cannot commit incest.